Browner still out; Clemons, Avril practice

RENTON, Wash. -- Seattle Seahawks defensive ends Chris Clemons and Cliff Avril returned to full participation at practice Thursday, but starting cornerback Brandon Browner still did not practice because of a hamstring injury.

Browner missed the opener last week against Carolina. The Seahawks could use his 6-foot-4 frame on Sunday night to help cover San Francisco 49ers veteran Anquan Boldin, one of the strongest wide receivers in the NFL. Boldin had 13 receptions for 208 yards in a season-opening victory over Green Bay.

It appears likely that Avril will play for the first time in Seahawks uniform. Avril came from Detroit as a free agent and was Seattle’s top offseason acquisition on defense, the man who coaches hoped could improve the team’s pass rush. But he missed the preseason with foot and hamstring injuries and did not play at Carolina.

Clemons, whose 11.5 sacks last season led the Seahawks, is recovering from offseason ACL surgery. His status for Sunday is unknown, but it’s clear he will return soon.

Defensive tackle Brandon Mebane was a limited participant in practice Thursday after missing practice Wednesday with an ankle injury. Defensive tackle Tony McDaniel did not practice because of a groin injury. Backup safety Jeron Johnson has not practiced this week after injuring a hamstring in the season opener.