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Marcell Dareus, Sammy Watkins headline Bills' non-participation lists

Marcell Dareus is on the Bills' active/non-football illness list because of an undisclosed medical issue. Al Messerschmidt/Getty Images

PITTSFORD, N.Y. -- The Buffalo Bills placed 10 players on non-participation lists as they opened training camp Friday at St. John Fisher College.

Each player is eligible to come off the lists at any time once he is cleared to practice by the medical staff. Once a player is taken off their respective active lists, they are not eligible to be transferred to a reserve list to start the regular season.

Here is a rundown of all 10 players, all of whom still count against the 90-man roster limit:

DT Marcell Dareus: The Bills' Pro Bowl defensive tackle was placed on the active/non-football illness list with an undisclosed medical issue. Dareus said he passed the Bills' conditioning test Friday. General manager Doug Whaley said Friday that Dareus' ailment is "nothing major."

WR Sammy Watkins: He begins camp on the active/physically unable to perform list, which isn't a surprise as the team continues to be cautious with his broken foot. It's still unclear how much Watkins will participate in training camp and the preseason, but the team's goal remains to have him ready for the regular-season opener.

RB Karlos Williams: Williams' conditioning was a hot topic during minicamp when he showed up overweight. Whaley said Friday that Williams needs to be "cleared medically" before he can work on getting "in condition to play football," although Whaley didn't offer specifics about Williams' medical issue. Ryan added, "We're not going to be putting a player at risk, through running conditioning tests or whatever it may be, if we don't think that that player is ready to take on that kind of challenge," hinting that Williams did not take part in a full conditioning test Friday.

DT Kyle Williams: As expected, the Bills' longest-tenured player begins camp on the active/physically unable to perform list after undergoing season-ending knee surgery late last season. Whaley said the team's priority is to get Williams healthy and ready for Week 1. That might mean very limited practice time for the Pro Bowl defensive tackle this summer, which isn't a big deal -- the veteran knows his stuff.

OLB Shaq Lawson: As expected, Lawson will begin camp -- and likely spend all of camp -- on the active/physically unable to perform list after undergoing shoulder surgery in May. Whaley kept open the possibility of the rookie first-round pick landing on the reserve/PUP list to start the season, which would keep him out at least six games, but added that all the reports on Lawson's recovery so far have been positive.

OLB Manny Lawson: The veteran was placed on the active/non-football injury list. Whaley said Friday that Lawson suffered an upper body injury while training shortly before reporting to camp, and that the team's medical staff was still in the process of offering a prognosis. Whaley added that he doesn't anticipate it to be a long-term injury, but "you never know." With Lawson also sidelined, IK Enemkpali is expected to get first-team reps at outside linebacker in Lawson's place.

WR Kolby Listenbee: The rookie sixth-round pick was placed on the active/non-football injury list as he continues to recover from a sports hernia that predates his joining the Bills.

WR Marcus Easley: Easley is still recovering from a serious knee injury suffered in Week 16 last season. He was placed on the active/physically unable to perform list.

OT Seantrel Henderson: Henderson missed the final four games of last season after he was diagnosed with Crohn's disease. He was placed on the active/non-football illness list, but Ryan said Henderson passed his conditioning test Friday.

CB Jonathan Dowling: He spent the final games of last season on the Bills' practice squad and is now on the active/non-football illness list with an undisclosed medical issue.